01 / Morfologi

Ciri dan struktur

Saprobic on dead branches of Urtica dioica. Sexual morph Ascomata 100–130 high, 70–110 μm diam., solitary, scattered or in small groups, erumpent through host epidermis to superficial, conical to mammiform, dark brown to black, coriaceous, smooth, easily removed from the host substrate, ostiolate with minute papilla. Ostioles 50–70 μm diam., papillate, black, shiny, smooth. Peridium 25–50 μm wide, comprising two cell types, outer layer composed of small, thick-walled cells of textura angularis to textura globulosa, surface heavily pigmented termed as scleroplectenchyma, thinner at the apex, wide at sides, inner layer composed of subhyaline or light brown relatively thin-walled cells of textura angularis, cells near the base comparatively larger. Hamathecium comprising numerous, dense, 1.5–2 μm wide, filamentous, septate, cellular pseudoparaphyses, branched and anastomosing, embedded in gelatinous matrix. Asci 60–140 × 9–11 μm (\(\bar{x} \) = 104.5 × 10 μm, n = 20), 8-spored, bitunicate, fissitunicate, cylindrical, rounded at the apex, pedicellate, numerous, with ocular chamber. Ascospores 35–40 × 4–6 μm (\(\bar{x} \) = 38.3 × 5.2 μm, n = 20), overlapping 1–2-seriate, initially hyaline, becoming yellowish brown at maturity, long fusiform, (8–)9-septate, constricted at the septa, narrowly rounded at both ends, smooth-walled, lacking a mucilaginous sheath. Asexual morph Undetermined. Culture characteristics: Ascospores germinating on MEA within 24 h. Colonies growing on MEA reaching 2 cm diam. in one week at 16 °C. Mycelium superficial, surface smooth, irregular, slightly raised, edge crenate, velutinous, from above white to pale yellow, reverse yellow.

Sebagian uraian ditampilkan dalam bahasa sumber; terjemahannya belum tersedia atau belum lolos pemeriksaan.

Rujukan: [1]

Pengukuran mengikuti objek, spesimen atau kultur, dan metode yang disebutkan sumber. Baca uraian asli untuk konteks lengkap.

02 / Kehidupan

Habitat dan peran ekologis

Saprobic on dead branches of Urtica dioica. Sexual morph Ascomata 100–130 high, 70–110 μm diam., solitary, scattered or in small groups, erumpent through host epidermis to superficial, conical to mammiform, dark brown to black, coriaceous, smooth, easily removed from the host substrate, ostiolate with minute papilla. Ostioles 50–70 μm diam., papillate, black, shiny, smooth. Peridium 25–50 μm wide, comprising two cell types, outer layer composed of small, thick-walled cells of textura angularis to textura globulosa, surface heavily pigmented termed as scleroplectenchyma, thinner at the apex, wide at sides, inner layer composed of subhyaline or light brown relatively thin-walled cells of textura angularis, cells near the base comparatively larger. Hamathecium comprising numerous, dense, 1.5–2 μm wide, filamentous, septate, cellular pseudoparaphyses, branched and anastomosing, embedded in gelatinous matrix. Asci 60–140 × 9–11 μm (\(\bar{x} \) = 104.5 × 10 μm, n = 20), 8-spored, bitunicate, fissitunicate, cylindrical, rounded at the apex, pedicellate, numerous, with ocular chamber. Ascospores 35–40 × 4–6 μm (\(\bar{x} \) = 38.3 × 5.2 μm, n = 20), overlapping 1–2-seriate, initially hyaline, becoming yellowish brown at maturity, long fusiform, (8–)9-septate, constricted at the septa, narrowly rounded at both ends, smooth-walled, lacking a mucilaginous sheath.

Sebagian uraian ditampilkan dalam bahasa sumber; terjemahannya belum tersedia atau belum lolos pemeriksaan.

Rujukan: [1]

Habitat atau konteks temuan
Informasi belum tersedia pada sumber rilis ini.
Substrat
kayu
Peran atau konteks
Pengurai
Temukan hubungan serupa
03 / Siklus hidup

Reproduksi

Saprobic on dead branches of Urtica dioica. Sexual morph Ascomata 100–130 high, 70–110 μm diam., solitary, scattered or in small groups, erumpent through host epidermis to superficial, conical to mammiform, dark brown to black, coriaceous, smooth, easily removed from the host substrate, ostiolate with minute papilla. Ostioles 50–70 μm diam., papillate, black, shiny, smooth. Peridium 25–50 μm wide, comprising two cell types, outer layer composed of small, thick-walled cells of textura angularis to textura globulosa, surface heavily pigmented termed as scleroplectenchyma, thinner at the apex, wide at sides, inner layer composed of subhyaline or light brown relatively thin-walled cells of textura angularis, cells near the base comparatively larger.

Sebagian uraian ditampilkan dalam bahasa sumber; terjemahannya belum tersedia atau belum lolos pemeriksaan.

Rujukan: [1]

04 / Batas informasi

Catatan identitas dan cakupan

  • Nama diterima mengikuti Catalogue of Life Base2026-09-11; uraian mempertahankan konsep dan nama sumber aslinya.
  • Preserve original monograph treatment localities, hosts and specimen/substrate scope. Materials examined document specimens, not the full global range.
  • Impor dan terjemahan otomatis belum diperiksa editorial per profil. Struktur reproduktif yang disebutkan tidak membuktikan seluruh siklus hidup diketahui.
  • Pengukuran pada uraian mengikuti spesimen, kultur, dan metode sumber; angka tidak diperluas ke semua populasi.

Catatan distribusi dalam narasi mengikuti sumbernya. Dokumentasi spesimen atau foto mengikuti catatan sumber. Bukti wilayah Indonesia dan penilaian konservasi tidak disimpulkan dari gambar.

05 / Dokumentasi

Melihat dengan konteks

Leptosphaeria urticae (MFLU 18-0591, holotype). a–c Appearance of ascomata on host surface. d Section through an ascoma. e Peridium. f Pseudoparaphyses. g–i Asci. j–m Ascospores. n Spore germination on MEA after 24 h. o Culture from above and below. Scale bars a, b = 500 µm, c = 200 μm, d = 100 μm, e, g–i = 50 μm, j–m = 10 μm, f = 5 μm

Jenis
Pelat ilmiah
Kredit
Penulis dan sumber artikel: Phookamsak, Rungtiwa; Hyde, Kevin D.; Jeewon, Rajesh; Bhat, D. Jayarama; Jones, E. B. Gareth; Maharachchikumbura, Sajeewa S. N.; Raspé, Olivier; Karunarathna, Samantha C.; Wanasinghe, Dhanushka N.; Hongsanan, Sinang; Doilom, Mingkwan; Tennakoon, Danushka S.; Machado, Alexandre R.; Firmino, André L.; Ghosh, Aniket; Karunarathna, Anuruddha; Mešić, Armin; Dutta, Arun Kumar; Thongbai, Benjarong; Devadatha, Bandarupalli; Norphanphoun, Chada; Senwanna, Chanokned; Wei, Deping; Pem, Dhandevi; Ackah, Frank Kwekucher; Wang, Gen-Nuo; Jiang, Hong-Bo; Madrid, Hugo; Lee, Hyang Burm; Goonasekara, Ishani D.; Manawasinghe, Ishara S.; Kušan, Ivana; Cano, Josep; Gené, Josepa; Li, Junfu; Das, Kanad; Acharya, Krishnendu; Raj, K. N. Anil; Latha, K. P. Deepna; Chethana, K. W. Thilini; He, Mao-Qiang; Dueñas, Margarita; Jadan, Margita; Martín, María P.; Samarakoon, Milan C.; Dayarathne, Monika C.; Raza, Mubashar; Park, Myung Soo; Telleria, M. Teresa; Chaiwan, Napalai; Matočec, Neven; de Silva, Nimali I.; Pereira, Olinto L.; Singh, Paras Nath; Manimohan, Patinjareveettil; Uniyal, Priyanka; Shang, Qiu-Ju; Bhatt, Rajendra P.; Perera, Rekhani H.; Alvarenga, Renato Lúcio Mendes; Nogal-Prata, Sandra; Singh, Sanjay K.; Vadthanarat, Santhiti; Oh, Seung-Yoon; Huang, Shi-Ke; Rana, Shiwali; Konta, Sirinapa; Paloi, Soumitra; Jayasiri, Subashini C.; Jeon, Sun Jeong; Mehmood, Tahir; Gibertoni, Tatiana Baptista; Nguyen, Thuong T. T.; Singh, Upendra; Thiyagaraja, Vinodhini; Sarma, V. Venkateswara; Dong, Wei; Yu, Xian-Dong; Lu, Yong-Zhong; Lim, Young Woon; Chen, Yun; Tkalčec, Zdenko; Zhang, Zhi-Feng; Luo, Zong-Long; Daranagama, Dinushani A.; Thambugala, Kasun M.; Tibpromma, Saowaluck; Camporesi, Erio; Bulgakov, Timur S.; Dissanayake, Asha J.; Senanayake, Indunil C.; Dai, Dong Qin; Tang, Li-Zhou; Khan, Sehroon; Zhang, Huang; Promputtha, Itthayakorn; Cai, Lei; Chomnunti, Putarak; Zhao, Rui-Lin; Lumyong, Saisamorn; Boonmee, Saranyaphat; Wen, Ting-Chi; Mortimer, Peter E.; Xu, Jianchu
Lisensi
CC BY 4.0
Sumber media
Buka entri sumber

Gambar menunjukkan objek menurut identifikasi sumber, bukan verifikasi independen spesimen oleh Fungi. Tidak ada skala baru yang ditambahkan.

06 / Rujukan

Sumber profil

  1. [1] Fungal diversity notes 929–1035: taxonomic and phylogenetic contributions on genera and species of fungi — Leptosphaeria urticaeBagian sumber: publisher:10.1007/s13225-019-00421-w:paragraph:130
    Tanggal penghimpunan sumber: 2026-10-12
    Teks sumber: CC BY 4.0 · Phookamsak, Rungtiwa; Hyde, Kevin D.; Jeewon, Rajesh; Bhat, D. Jayarama; Jones, E. B. Gareth; Maharachchikumbura, Sajeewa S. N.; Raspé, Olivier; Karunarathna, Samantha C.; Wanasinghe, Dhanushka N.; Hongsanan, Sinang; Doilom, Mingkwan; Tennakoon, Danushka S.; Machado, Alexandre R.; Firmino, André L.; Ghosh, Aniket; Karunarathna, Anuruddha; Mešić, Armin; Dutta, Arun Kumar; Thongbai, Benjarong; Devadatha, Bandarupalli; Norphanphoun, Chada; Senwanna, Chanokned; Wei, Deping; Pem, Dhandevi; Ackah, Frank Kwekucher; Wang, Gen-Nuo; Jiang, Hong-Bo; Madrid, Hugo; Lee, Hyang Burm; Goonasekara, Ishani D.; Manawasinghe, Ishara S.; Kušan, Ivana; Cano, Josep; Gené, Josepa; Li, Junfu; Das, Kanad; Acharya, Krishnendu; Raj, K. N. Anil; Latha, K. P. Deepna; Chethana, K. W. Thilini; He, Mao-Qiang; Dueñas, Margarita; Jadan, Margita; Martín, María P.; Samarakoon, Milan C.; Dayarathne, Monika C.; Raza, Mubashar; Park, Myung Soo; Telleria, M. Teresa; Chaiwan, Napalai; Matočec, Neven; de Silva, Nimali I.; Pereira, Olinto L.; Singh, Paras Nath; Manimohan, Patinjareveettil; Uniyal, Priyanka; Shang, Qiu-Ju; Bhatt, Rajendra P.; Perera, Rekhani H.; Alvarenga, Renato Lúcio Mendes; Nogal-Prata, Sandra; Singh, Sanjay K.; Vadthanarat, Santhiti; Oh, Seung-Yoon; Huang, Shi-Ke; Rana, Shiwali; Konta, Sirinapa; Paloi, Soumitra; Jayasiri, Subashini C.; Jeon, Sun Jeong; Mehmood, Tahir; Gibertoni, Tatiana Baptista; Nguyen, Thuong T. T.; Singh, Upendra; Thiyagaraja, Vinodhini; Sarma, V. Venkateswara; Dong, Wei; Yu, Xian-Dong; Lu, Yong-Zhong; Lim, Young Woon; Chen, Yun; Tkalčec, Zdenko; Zhang, Zhi-Feng; Luo, Zong-Long; Daranagama, Dinushani A.; Thambugala, Kasun M.; Tibpromma, Saowaluck; Camporesi, Erio; Bulgakov, Timur S.; Dissanayake, Asha J.; Senanayake, Indunil C.; Dai, Dong Qin; Tang, Li-Zhou; Khan, Sehroon; Zhang, Huang; Promputtha, Itthayakorn; Cai, Lei; Chomnunti, Putarak; Zhao, Rui-Lin; Lumyong, Saisamorn; Boonmee, Saranyaphat; Wen, Ting-Chi; Mortimer, Peter E.; Xu, Jianchu
  2. [2] Catalogue of Life Base2026-09-11 — Leptosphaeria urticaeBagian sumber: ColDP NameUsage.tsv ID 6PK9P; accepted species; source 2073
    Tanggal penghimpunan sumber: 2026-10-12
  3. [3] Dokumentasi visual — Leptosphaeria urticaeBagian sumber: Individual media record; identification follows the source, not independent determination
    Tanggal penghimpunan sumber: 2026-10-12
    Media sumber: CC BY 4.0 · Penulis dan sumber artikel: Phookamsak, Rungtiwa; Hyde, Kevin D.; Jeewon, Rajesh; Bhat, D. Jayarama; Jones, E. B. Gareth; Maharachchikumbura, Sajeewa S. N.; Raspé, Olivier; Karunarathna, Samantha C.; Wanasinghe, Dhanushka N.; Hongsanan, Sinang; Doilom, Mingkwan; Tennakoon, Danushka S.; Machado, Alexandre R.; Firmino, André L.; Ghosh, Aniket; Karunarathna, Anuruddha; Mešić, Armin; Dutta, Arun Kumar; Thongbai, Benjarong; Devadatha, Bandarupalli; Norphanphoun, Chada; Senwanna, Chanokned; Wei, Deping; Pem, Dhandevi; Ackah, Frank Kwekucher; Wang, Gen-Nuo; Jiang, Hong-Bo; Madrid, Hugo; Lee, Hyang Burm; Goonasekara, Ishani D.; Manawasinghe, Ishara S.; Kušan, Ivana; Cano, Josep; Gené, Josepa; Li, Junfu; Das, Kanad; Acharya, Krishnendu; Raj, K. N. Anil; Latha, K. P. Deepna; Chethana, K. W. Thilini; He, Mao-Qiang; Dueñas, Margarita; Jadan, Margita; Martín, María P.; Samarakoon, Milan C.; Dayarathne, Monika C.; Raza, Mubashar; Park, Myung Soo; Telleria, M. Teresa; Chaiwan, Napalai; Matočec, Neven; de Silva, Nimali I.; Pereira, Olinto L.; Singh, Paras Nath; Manimohan, Patinjareveettil; Uniyal, Priyanka; Shang, Qiu-Ju; Bhatt, Rajendra P.; Perera, Rekhani H.; Alvarenga, Renato Lúcio Mendes; Nogal-Prata, Sandra; Singh, Sanjay K.; Vadthanarat, Santhiti; Oh, Seung-Yoon; Huang, Shi-Ke; Rana, Shiwali; Konta, Sirinapa; Paloi, Soumitra; Jayasiri, Subashini C.; Jeon, Sun Jeong; Mehmood, Tahir; Gibertoni, Tatiana Baptista; Nguyen, Thuong T. T.; Singh, Upendra; Thiyagaraja, Vinodhini; Sarma, V. Venkateswara; Dong, Wei; Yu, Xian-Dong; Lu, Yong-Zhong; Lim, Young Woon; Chen, Yun; Tkalčec, Zdenko; Zhang, Zhi-Feng; Luo, Zong-Long; Daranagama, Dinushani A.; Thambugala, Kasun M.; Tibpromma, Saowaluck; Camporesi, Erio; Bulgakov, Timur S.; Dissanayake, Asha J.; Senanayake, Indunil C.; Dai, Dong Qin; Tang, Li-Zhou; Khan, Sehroon; Zhang, Huang; Promputtha, Itthayakorn; Cai, Lei; Chomnunti, Putarak; Zhao, Rui-Lin; Lumyong, Saisamorn; Boonmee, Saranyaphat; Wen, Ting-Chi; Mortimer, Peter E.; Xu, Jianchu

Adaptasi teks: CC BY 4.0. Phookamsak, Rungtiwa; Hyde, Kevin D.; Jeewon, Rajesh; Bhat, D. Jayarama; Jones, E. B. Gareth; Maharachchikumbura, Sajeewa S. N.; Raspé, Olivier; Karunarathna, Samantha C.; Wanasinghe, Dhanushka N.; Hongsanan, Sinang; Doilom, Mingkwan; Tennakoon, Danushka S.; Machado, Alexandre R.; Firmino, André L.; Ghosh, Aniket; Karunarathna, Anuruddha; Mešić, Armin; Dutta, Arun Kumar; Thongbai, Benjarong; Devadatha, Bandarupalli; Norphanphoun, Chada; Senwanna, Chanokned; Wei, Deping; Pem, Dhandevi; Ackah, Frank Kwekucher; Wang, Gen-Nuo; Jiang, Hong-Bo; Madrid, Hugo; Lee, Hyang Burm; Goonasekara, Ishani D.; Manawasinghe, Ishara S.; Kušan, Ivana; Cano, Josep; Gené, Josepa; Li, Junfu; Das, Kanad; Acharya, Krishnendu; Raj, K. N. Anil; Latha, K. P. Deepna; Chethana, K. W. Thilini; He, Mao-Qiang; Dueñas, Margarita; Jadan, Margita; Martín, María P.; Samarakoon, Milan C.; Dayarathne, Monika C.; Raza, Mubashar; Park, Myung Soo; Telleria, M. Teresa; Chaiwan, Napalai; Matočec, Neven; de Silva, Nimali I.; Pereira, Olinto L.; Singh, Paras Nath; Manimohan, Patinjareveettil; Uniyal, Priyanka; Shang, Qiu-Ju; Bhatt, Rajendra P.; Perera, Rekhani H.; Alvarenga, Renato Lúcio Mendes; Nogal-Prata, Sandra; Singh, Sanjay K.; Vadthanarat, Santhiti; Oh, Seung-Yoon; Huang, Shi-Ke; Rana, Shiwali; Konta, Sirinapa; Paloi, Soumitra; Jayasiri, Subashini C.; Jeon, Sun Jeong; Mehmood, Tahir; Gibertoni, Tatiana Baptista; Nguyen, Thuong T. T.; Singh, Upendra; Thiyagaraja, Vinodhini; Sarma, V. Venkateswara; Dong, Wei; Yu, Xian-Dong; Lu, Yong-Zhong; Lim, Young Woon; Chen, Yun; Tkalčec, Zdenko; Zhang, Zhi-Feng; Luo, Zong-Long; Daranagama, Dinushani A.; Thambugala, Kasun M.; Tibpromma, Saowaluck; Camporesi, Erio; Bulgakov, Timur S.; Dissanayake, Asha J.; Senanayake, Indunil C.; Dai, Dong Qin; Tang, Li-Zhou; Khan, Sehroon; Zhang, Huang; Promputtha, Itthayakorn; Cai, Lei; Chomnunti, Putarak; Zhao, Rui-Lin; Lumyong, Saisamorn; Boonmee, Saranyaphat; Wen, Ting-Chi; Mortimer, Peter E.; Xu, Jianchu — Fungal diversity notes 929–1035: taxonomic and phylogenetic contributions on genera and species of fungi — Leptosphaeria urticae

Uraian asli dan riwayat sumber

Riwayat atau halaman sumber

Ringkasan sumber

Saprobic on dead branches of Urtica dioica. Sexual morph Ascomata 100–130 high, 70–110 μm diam., solitary, scattered or in small groups, erumpent through host epidermis to superficial, conical to mammiform, dark brown to black, coriaceous, smooth, easily removed from the host substrate, ostiolate with minute papilla. Ostioles 50–70 μm diam., papillate, black, shiny, smooth. Peridium 25–50 μm wide, comprising two cell types, outer layer composed of small, thick-walled cells of textura angularis to textura globulosa, surface heavily pigmented termed as scleroplectenchyma, thinner at the apex, wide at sides, inner layer composed of subhyaline or light brown relatively thin-walled cells of textura angularis, cells near the base comparatively larger.

Revisi sumber

Morfologi sumber

Saprobic on dead branches of Urtica dioica. Sexual morph Ascomata 100–130 high, 70–110 μm diam., solitary, scattered or in small groups, erumpent through host epidermis to superficial, conical to mammiform, dark brown to black, coriaceous, smooth, easily removed from the host substrate, ostiolate with minute papilla. Ostioles 50–70 μm diam., papillate, black, shiny, smooth. Peridium 25–50 μm wide, comprising two cell types, outer layer composed of small, thick-walled cells of textura angularis to textura globulosa, surface heavily pigmented termed as scleroplectenchyma, thinner at the apex, wide at sides, inner layer composed of subhyaline or light brown relatively thin-walled cells of textura angularis, cells near the base comparatively larger. Hamathecium comprising numerous, dense, 1.5–2 μm wide, filamentous, septate, cellular pseudoparaphyses, branched and anastomosing, embedded in gelatinous matrix. Asci 60–140 × 9–11 μm (\(\bar{x} \) = 104.5 × 10 μm, n = 20), 8-spored, bitunicate, fissitunicate, cylindrical, rounded at the apex, pedicellate, numerous, with ocular chamber. Ascospores 35–40 × 4–6 μm (\(\bar{x} \) = 38.3 × 5.2 μm, n = 20), overlapping 1–2-seriate, initially hyaline, becoming yellowish brown at maturity, long fusiform, (8–)9-septate, constricted at the septa, narrowly rounded at both ends, smooth-walled, lacking a mucilaginous sheath. Asexual morph Undetermined. Culture characteristics: Ascospores germinating on MEA within 24 h. Colonies growing on MEA reaching 2 cm diam. in one week at 16 °C. Mycelium superficial, surface smooth, irregular, slightly raised, edge crenate, velutinous, from above white to pale yellow, reverse yellow.

Revisi sumber

Ekologi sumber

Saprobic on dead branches of Urtica dioica. Sexual morph Ascomata 100–130 high, 70–110 μm diam., solitary, scattered or in small groups, erumpent through host epidermis to superficial, conical to mammiform, dark brown to black, coriaceous, smooth, easily removed from the host substrate, ostiolate with minute papilla. Ostioles 50–70 μm diam., papillate, black, shiny, smooth. Peridium 25–50 μm wide, comprising two cell types, outer layer composed of small, thick-walled cells of textura angularis to textura globulosa, surface heavily pigmented termed as scleroplectenchyma, thinner at the apex, wide at sides, inner layer composed of subhyaline or light brown relatively thin-walled cells of textura angularis, cells near the base comparatively larger. Hamathecium comprising numerous, dense, 1.5–2 μm wide, filamentous, septate, cellular pseudoparaphyses, branched and anastomosing, embedded in gelatinous matrix. Asci 60–140 × 9–11 μm (\(\bar{x} \) = 104.5 × 10 μm, n = 20), 8-spored, bitunicate, fissitunicate, cylindrical, rounded at the apex, pedicellate, numerous, with ocular chamber. Ascospores 35–40 × 4–6 μm (\(\bar{x} \) = 38.3 × 5.2 μm, n = 20), overlapping 1–2-seriate, initially hyaline, becoming yellowish brown at maturity, long fusiform, (8–)9-septate, constricted at the septa, narrowly rounded at both ends, smooth-walled, lacking a mucilaginous sheath.

Revisi sumber

Struktur reproduksi sumber

Saprobic on dead branches of Urtica dioica. Sexual morph Ascomata 100–130 high, 70–110 μm diam., solitary, scattered or in small groups, erumpent through host epidermis to superficial, conical to mammiform, dark brown to black, coriaceous, smooth, easily removed from the host substrate, ostiolate with minute papilla. Ostioles 50–70 μm diam., papillate, black, shiny, smooth. Peridium 25–50 μm wide, comprising two cell types, outer layer composed of small, thick-walled cells of textura angularis to textura globulosa, surface heavily pigmented termed as scleroplectenchyma, thinner at the apex, wide at sides, inner layer composed of subhyaline or light brown relatively thin-walled cells of textura angularis, cells near the base comparatively larger.

Revisi sumber

Rujukan halaman ini

Fungi. Leptosphaeria urticae. Rilis data fungi-2026-10-12-expansion-2. https://fungi-atlas.mgadib.chatgpt.site/jamur/leptosphaeria-urticae/.

Status: impor sumber; pemeriksaan editorial per profil belum selesai. Untuk klaim biologis, gunakan rujukan pada bagian terkait dan periksa konteks sumber aslinya.